Topography and planting density affect understory plant diversity in conifer plantations through development of stand canopy closure.

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Title: Topography and planting density affect understory plant diversity in conifer plantations through development of stand canopy closure.
Authors: Matsushita, Michinari1 (AUTHOR) matsushita@affrc.go.jp, Takashima, Yuya1 (AUTHOR), Tamura, Akira1 (AUTHOR)
Source: Canadian Journal of Forest Research. 11/20/2025, Vol. 55, p1-10. 10p.
Subject Terms: *Biodiversity, *Botany, *Forest density, *Species diversity, *Reforestation, Topography, Coniferous forests, Plant spacing
Abstract: Sustainably managed plantations not only provide high-quality timber but also contribute to biodiversity. Clarifying the direct and indirect effects of planting density and topography on the development and diversity of understory vegetation is important from the perspective of the diversity-retaining function of young plantations. This study conducted a vegetation survey in plantation stands of Japanese cedar, 4–7 years after planting. These plantations varied in initial planting density (ranging 1000–3000 trees/ha), and contained topographic variations from ridges to lower slopes. The direct and indirect effects of initial planting density and topography on species composition, diversity, and height growth of understory vegetation were investigated. Topographic variation affected stand canopy closure and the mean height growth rates of planted cedars. Topography effect was directly related to vegetation height. High initial planting density accelerated stand canopy closure through crown development of planted trees. Through stand canopy closure, initial planting density and topography indirectly affected plant diversity, although their direct effects were not significantly detected. Initial planting density and topography impacted vegetation diversity mainly through the degree of canopy closure, rather than directly. These findings are useful when planning reforestation schedules and spatial designs within the landscape while taking diversity into consideration. [ABSTRACT FROM AUTHOR]
Copyright of Canadian Journal of Forest Research is the property of Canadian Science Publishing and its content may not be copied or emailed to multiple sites without the copyright holder's express written permission. Additionally, content may not be used with any artificial intelligence tools or machine learning technologies. However, users may print, download, or email articles for individual use. This abstract may be abridged. No warranty is given about the accuracy of the copy. Users should refer to the original published version of the material for the full abstract. (Copyright applies to all Abstracts.)
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Items – Name: Title
  Label: Title
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  Data: Topography and planting density affect understory plant diversity in conifer plantations through development of stand canopy closure.
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  Data: <searchLink fieldCode="AR" term="%22Matsushita%2C+Michinari%22">Matsushita, Michinari</searchLink><relatesTo>1</relatesTo> (AUTHOR)<i> matsushita@affrc.go.jp</i><br /><searchLink fieldCode="AR" term="%22Takashima%2C+Yuya%22">Takashima, Yuya</searchLink><relatesTo>1</relatesTo> (AUTHOR)<br /><searchLink fieldCode="AR" term="%22Tamura%2C+Akira%22">Tamura, Akira</searchLink><relatesTo>1</relatesTo> (AUTHOR)
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  Data: <searchLink fieldCode="JN" term="%22Canadian+Journal+of+Forest+Research%22">Canadian Journal of Forest Research</searchLink>. 11/20/2025, Vol. 55, p1-10. 10p.
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  Data: *<searchLink fieldCode="DE" term="%22Biodiversity%22">Biodiversity</searchLink><br />*<searchLink fieldCode="DE" term="%22Botany%22">Botany</searchLink><br />*<searchLink fieldCode="DE" term="%22Forest+density%22">Forest density</searchLink><br />*<searchLink fieldCode="DE" term="%22Species+diversity%22">Species diversity</searchLink><br />*<searchLink fieldCode="DE" term="%22Reforestation%22">Reforestation</searchLink><br /><searchLink fieldCode="DE" term="%22Topography%22">Topography</searchLink><br /><searchLink fieldCode="DE" term="%22Coniferous+forests%22">Coniferous forests</searchLink><br /><searchLink fieldCode="DE" term="%22Plant+spacing%22">Plant spacing</searchLink>
– Name: Abstract
  Label: Abstract
  Group: Ab
  Data: Sustainably managed plantations not only provide high-quality timber but also contribute to biodiversity. Clarifying the direct and indirect effects of planting density and topography on the development and diversity of understory vegetation is important from the perspective of the diversity-retaining function of young plantations. This study conducted a vegetation survey in plantation stands of Japanese cedar, 4–7 years after planting. These plantations varied in initial planting density (ranging 1000–3000 trees/ha), and contained topographic variations from ridges to lower slopes. The direct and indirect effects of initial planting density and topography on species composition, diversity, and height growth of understory vegetation were investigated. Topographic variation affected stand canopy closure and the mean height growth rates of planted cedars. Topography effect was directly related to vegetation height. High initial planting density accelerated stand canopy closure through crown development of planted trees. Through stand canopy closure, initial planting density and topography indirectly affected plant diversity, although their direct effects were not significantly detected. Initial planting density and topography impacted vegetation diversity mainly through the degree of canopy closure, rather than directly. These findings are useful when planning reforestation schedules and spatial designs within the landscape while taking diversity into consideration. [ABSTRACT FROM AUTHOR]
– Name: AbstractSuppliedCopyright
  Label:
  Group: Ab
  Data: <i>Copyright of Canadian Journal of Forest Research is the property of Canadian Science Publishing and its content may not be copied or emailed to multiple sites without the copyright holder's express written permission. Additionally, content may not be used with any artificial intelligence tools or machine learning technologies. However, users may print, download, or email articles for individual use. This abstract may be abridged. No warranty is given about the accuracy of the copy. Users should refer to the original published version of the material for the full abstract.</i> (Copyright applies to all Abstracts.)
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RecordInfo BibRecord:
  BibEntity:
    Identifiers:
      – Type: doi
        Value: 10.1139/cjfr-2025-0080
    Languages:
      – Code: eng
        Text: English
    PhysicalDescription:
      Pagination:
        PageCount: 10
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    Subjects:
      – SubjectFull: Biodiversity
        Type: general
      – SubjectFull: Botany
        Type: general
      – SubjectFull: Forest density
        Type: general
      – SubjectFull: Species diversity
        Type: general
      – SubjectFull: Reforestation
        Type: general
      – SubjectFull: Topography
        Type: general
      – SubjectFull: Coniferous forests
        Type: general
      – SubjectFull: Plant spacing
        Type: general
    Titles:
      – TitleFull: Topography and planting density affect understory plant diversity in conifer plantations through development of stand canopy closure.
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          Name:
            NameFull: Matsushita, Michinari
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            NameFull: Takashima, Yuya
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            NameFull: Tamura, Akira
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            – D: 20
              M: 11
              Text: 11/20/2025
              Type: published
              Y: 2025
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              Value: 55
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            – TitleFull: Canadian Journal of Forest Research
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