A phosphorelay circuit drives extracellular alkalinization in receptor kinase-mediated immune and cell-wall damage signaling. - Zhai, Keran; Derbyshire, Paul; Zhang, Songyuan; Choi, Sera; Wang, Limin; Song, Beibei; Kinoshita, Toshinori; Zhou, Jian-Min; Menke, Frank L.H.; Bender, Kyle W.; Zipfel, Cyril (Molecular Cell, 2026)