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Biological Validation and Agnostic Experiments for Extant and Extinct Microbial Life within the Martian Subsurface.

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Title: Biological Validation and Agnostic Experiments for Extant and Extinct Microbial Life within the Martian Subsurface.
Authors: Perl SM; Earth, Planetary, and Space Sciences, University of California, Los Angeles, California, USA.; Mineral Sciences, Los Angeles Natural History Museum, Los Angeles, California, USA.; Blue Marble Space Institute of Science, Seattle, Washington, USA., Cockell CS; UK Centre for Astrobiology, University of Edinburgh, Edinburgh, UK.; Edinburgh Centre for Planetary Sciences, University of Edinburgh, Edinburgh, UK., Fischer WW; Geological and Planetary Sciences, California Institute of Technology, Pasadena, California, USA., Sharma S; Carnegie Institute of Science, Washington, District of Columbia, USA., Lloyd KG; Department of Earth Sciences, University of Southern California, Los Angeles, California, USA., Wilhelm MB; NASA Ames Research Center, Mountain View, California, USA., Corsetti FA; Department of Earth Sciences, University of Southern California, Los Angeles, California, USA., Treude T; Earth, Planetary, and Space Sciences, University of California, Los Angeles, California, USA., Leadbetter J; Geological and Planetary Sciences, California Institute of Technology, Pasadena, California, USA., Giovannelli D; University of Naples Federico II, Naples, Italy., Foreman CM; Chemical & Biological Engineering, Montana State University, Bozeman, Montana, USA., Freedman K; Department of Bioengineering, University of California, Riverside, Riverside, California, USA., Lynch K; NASA Headquarters, Washington, District of Columbia, USA., Homann M; Amentum Clean Energy Ltd, Oxfordshire, UK., Som SM; Mineral Sciences, Los Angeles Natural History Museum, Los Angeles, California, USA., Wilson L; School of Physics, Engineering and Technology, York University, York, UK., Karunatillake S; Department of Geology and Geophysics, Louisiana State University, Baton Rouge, Louisiana, USA., Drake H; Department of Biology and Environmental Science, Linnaeus University, Växjö, Sweden., Bywaters K; Honeybee Robotics, Altadena, California, USA., Mikucki J; Department of Microbiology, University of Tennessee, Knoxville, Tennessee, USA., Celestian AJ; Blue Marble Space Institute of Science, Seattle, Washington, USA., Chen P; NASA Jet Propulsion Laboratory, Pasadena, California, USA., Guan L; NASA Jet Propulsion Laboratory, Pasadena, California, USA., Ibarra Y; College of Science and Engineering, San Francisco State University, San Francisco, California, USA., Boland J; NASA Jet Propulsion Laboratory, Pasadena, California, USA., Hosseini S; NASA Jet Propulsion Laboratory, Pasadena, California, USA., Tosi PL; Kiln Energy, Los Angeles, California, USA., Nisson DM; NASA Ames Research Center, Mountain View, California, USA., Murphy A; Planetary Science Institute, Tucson, Arizona, USA., Garner M; Chemical & Biological Engineering, Montana State University, Bozeman, Montana, USA., Thei F; Elements SRL, Cesena, Italy., Biondi M; Elements SRL, Cesena, Italy., Propster P; California Institute of Technology, Pasadena, California, USA.
Source: Astrobiology [Astrobiology] 2026 Jul; Vol. 26 (1_suppl), pp. 156S-171S. Date of Electronic Publication: 2026 Jul 15.
Publication Type: Journal Article; Review
Journal Info: Publisher: Mary Ann Liebert, Inc Country of Publication: United States NLM ID: 101088083 Publication Model: Print-Electronic Cited Medium: Internet ISSN: 1557-8070 (Electronic) Linking ISSN: 15578070 NLM ISO Abbreviation: Astrobiology Subsets: MEDLINE
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NASA Ames Research Center, Mountain View, California, USA.<br /><searchLink fieldCode="AU" term="%22Corsetti+FA%22">Corsetti FA</searchLink>; Department of Earth Sciences, University of Southern California, Los Angeles, California, USA.<br /><searchLink fieldCode="AU" term="%22Treude+T%22">Treude T</searchLink>; Earth, Planetary, and Space Sciences, University of California, Los Angeles, California, USA.<br /><searchLink fieldCode="AU" term="%22Leadbetter+J%22">Leadbetter J</searchLink>; Geological and Planetary Sciences, California Institute of Technology, Pasadena, California, USA.<br /><searchLink fieldCode="AU" term="%22Giovannelli+D%22">Giovannelli D</searchLink>; University of Naples Federico II, Naples, Italy.<br /><searchLink fieldCode="AU" term="%22Foreman+CM%22">Foreman CM</searchLink>; Chemical & Biological Engineering, Montana State University, Bozeman, Montana, USA.<br /><searchLink fieldCode="AU" term="%22Freedman+K%22">Freedman K</searchLink>; Department of Bioengineering, University of California, Riverside, Riverside, California, USA.<br /><searchLink fieldCode="AU" term="%22Lynch+K%22">Lynch K</searchLink>; NASA Headquarters, Washington, District of Columbia, USA.<br /><searchLink fieldCode="AU" term="%22Homann+M%22">Homann M</searchLink>; Amentum Clean Energy Ltd, Oxfordshire, UK.<br /><searchLink fieldCode="AU" term="%22Som+SM%22">Som SM</searchLink>; Mineral Sciences, Los Angeles Natural History Museum, Los Angeles, California, USA.<br /><searchLink fieldCode="AU" term="%22Wilson+L%22">Wilson L</searchLink>; School of Physics, Engineering and Technology, York University, York, UK.<br /><searchLink fieldCode="AU" term="%22Karunatillake+S%22">Karunatillake S</searchLink>; Department of Geology and Geophysics, Louisiana State University, Baton Rouge, Louisiana, USA.<br /><searchLink fieldCode="AU" term="%22Drake+H%22">Drake H</searchLink>; Department of Biology and Environmental Science, Linnaeus University, Växjö, Sweden.<br /><searchLink fieldCode="AU" term="%22Bywaters+K%22">Bywaters K</searchLink>; Honeybee Robotics, Altadena, California, USA.<br /><searchLink fieldCode="AU" term="%22Mikucki+J%22">Mikucki J</searchLink>; Department of Microbiology, University of Tennessee, Knoxville, Tennessee, USA.<br /><searchLink fieldCode="AU" term="%22Celestian+AJ%22">Celestian AJ</searchLink>; Blue Marble Space Institute of Science, Seattle, Washington, USA.<br /><searchLink fieldCode="AU" term="%22Chen+P%22">Chen P</searchLink>; NASA Jet Propulsion Laboratory, Pasadena, California, USA.<br /><searchLink fieldCode="AU" term="%22Guan+L%22">Guan L</searchLink>; NASA Jet Propulsion Laboratory, Pasadena, California, USA.<br /><searchLink fieldCode="AU" term="%22Ibarra+Y%22">Ibarra Y</searchLink>; College of Science and Engineering, San Francisco State University, San Francisco, California, USA.<br /><searchLink fieldCode="AU" term="%22Boland+J%22">Boland J</searchLink>; NASA Jet Propulsion Laboratory, Pasadena, California, USA.<br /><searchLink fieldCode="AU" term="%22Hosseini+S%22">Hosseini S</searchLink>; NASA Jet Propulsion Laboratory, Pasadena, California, USA.<br /><searchLink fieldCode="AU" term="%22Tosi+PL%22">Tosi PL</searchLink>; Kiln Energy, Los Angeles, California, USA.<br /><searchLink fieldCode="AU" term="%22Nisson+DM%22">Nisson DM</searchLink>; NASA Ames Research Center, Mountain View, California, USA.<br /><searchLink fieldCode="AU" term="%22Murphy+A%22">Murphy A</searchLink>; Planetary Science Institute, Tucson, Arizona, USA.<br /><searchLink fieldCode="AU" term="%22Garner+M%22">Garner M</searchLink>; Chemical & Biological Engineering, Montana State University, Bozeman, Montana, USA.<br /><searchLink fieldCode="AU" term="%22Thei+F%22">Thei F</searchLink>; Elements SRL, Cesena, Italy.<br /><searchLink fieldCode="AU" term="%22Biondi+M%22">Biondi M</searchLink>; Elements SRL, Cesena, Italy.<br /><searchLink fieldCode="AU" term="%22Propster+P%22">Propster P</searchLink>; California Institute of Technology, Pasadena, California, USA.
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